Developmental checkpoints and feedback circuits time insect maturation

Kim Furbo Rewitz, Naoki Yamanaka, Michael B. O'Connor

81 Citations (Scopus)

Abstract

The transition from juvenile to adult is a fundamental process that allows animals to allocate resource toward reproduction after completing a certain amount of growth. In insects, growth to a species-specific target size induces pulses of the steroid hormone ecdysone that triggers metamorphosis and reproductive maturation. The past few years have seen significant progress in understanding the interplay of mechanisms that coordinate timing of ecdysone production and release. These studies show that the neuroendocrine system monitors complex size-related and nutritional signals, as well as external cues, to time production and release of ecdysone. Based on results discussed here, we suggest that developmental progression to adulthood is controlled by checkpoints that regulate the genetic timing program enabling it to adapt to different environmental conditions. These checkpoints utilize a number of signaling pathways to modulate ecdysone production in the prothoracic gland. Release of ecdysone activates an autonomous cascade of both feedforward and feedback signals that determine the duration of the ecdysone pulse at each developmental transitions. Conservation of the genetic mechanisms that coordinate the juvenile-adult transition suggests that insights from the fruit fly Drosophila will provide a framework for future investigation of developmental timing in metazoans.

Original languageEnglish
Title of host publicationAnimal metamorphosis
EditorsYun-Bo Shi
Number of pages33
Volume103
PublisherAcademic Press
Publication date2013
Pages1-33
Chapter1
ISBN (Print)978-0-12-385979-2
DOIs
Publication statusPublished - 2013
SeriesCurrent Topics in Developmental Biology
Volume103
ISSN0070-2153

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