Structure of the AvrBs3-DNA complex provides new insights into the initial thymine-recognition mechanism

Stefano Stella, Rafael Molina, Igor Yefimenko, Jesús Prieto, George Silva, Claudia Bertonati, Alexandre Juillerat, Phillippe Duchateau, Guillermo Montoya*

*Corresponding author af dette arbejde
25 Citationer (Scopus)

Abstract

Transcription activator-like effectors contain a DNA-binding domain organized in tandem repeats. The repeats include two adjacent residues known as the repeat variable di-residue, which recognize a single base pair, establishing a direct code between the dipeptides and the target DNA. This feature suggests this scaffold as an excellent candidate to generate new protein-DNA specificities for biotechnological applications. Here, the crystal structure of AvrBs3 (residues 152-895, molecular mass 82kDa) in complex with its target DNA sequence is presented, revealing a new mode of interaction with the initial thymine of the target sequence, together with an analysis of both the binding specificity and the thermodynamic properties of AvrBs3. This study quantifies the affinity and the specificity between AvrBs3 and its target DNA. Moreover, in vitro and in vivo analyses reveal that AvrBs3 does not show a strict nucleotide-binding preference for the nucleotide at the zero position of the DNA, widening the number of possible sequences that could be targeted by this scaffold.

OriginalsprogEngelsk
TidsskriftActa Crystallographica Section D: Biological Crystallography
Vol/bind69
Udgave nummer9
Sider (fra-til)1707-1716
Antal sider10
ISSN0907-4449
DOI
StatusUdgivet - 1 sep. 2013

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